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Sternberg, C. H., 1922.

Explorations of the Permian of Texas and the chalk of Kansas, 1918.

Kansas Academy of Science, Transactions  30(1):119-120.

 

 

Copyright © 2002-2013 by Mike Everhart

Created 08/28//2002 -Last updated 02/17/2013

 

 

 

 

LEFT: The specimen of Tylosaurus proriger (USNM 8898) found by George Sternberg in 1918; On exhibit in the United States National Museum (Smithsonian).

LEFT: In the National Geographic IMAX movie Sea Monsters (2007), there is a Tylosaurus lurking somewhere under the surface, waiting for its next meal including a small polycotylid plesiosaur (Dolichorhynchops osborni) shown overhead. While not entirely correct scientifically regarding the behavior of this mosasaur, the movie itself is based on the actual 1918 discovery of a Tylosaurus proriger skeleton with the bones of a juvenile plesiosaur inside the rib cage. The discovery was made by Charles H. Sternberg in Logan County, Kansas, and the specimen was sold to the United States National Museum (USNM - Smithsonian) where it has been on display since about 1920. 

Although Sternberg published the discovery, it was more or less lost in history until I came across the report in his two page note in the Transactions of the Kansas Academy of Science (1922):

“…I joined them in the Kansas chalk in Logan county, on Butte creek. I was so fortunate as to find a fine tylosaur skeleton the second day in the field. There were twenty-one feet of the skeleton present in fine chalk. The complete skull was crushed laterally. Nearly the complete front arches and limbs were present, as was also the pelvic bones and both femora. All the vertebrae to well into the caudal region beyond the lateral spines were continuous, with the ribs in the dorsal region. Between the ribs was a large part of a huge plesiosaur with many half-digested bones, including the large humeri part of the coracoscapula, phalanges, vertebrae, and, strangest of all, the stomach stones, showing that this huge tylosaur, that was about twenty-nine feet long, had swallowed this plesiosaur in large enough chunks to include the stomach. How powerful the gastric juice that could dissolve these big bones! This specimen I sent to the United States National Museum.”

gilmorec.jpg (16345 bytes) Wherein Charles Sternberg describes the important discovery of a plesiosaur found as stomach contents of a large Tylosaurus proriger mosasaur, and the collection of a very rare mosasaur that eventually will be named after him ("Clidastes sternbergi"; now Eonatator (Halisaurus) sternbergi). The Tylosaurus and plesiosaur specimens were acquired by the United States National Museum (Smithsonian) where the mosasaur (LEFT) was placed on exhibit.  The picture above was taken in 2001.  Charles Gilmore (1921) described the mosasaur specimen in a brief article in Scientific American, but neglected to mention the plesiosaur even though he discussed the eating habits of mosasaurs in general. When examined in 2001, the plesiosaur specimen also included about a dozen small stones (gastroliths), a shark tooth (Squalicorax falcatus) and part of a large fish vertebrae (Xiphactinus audax).                                                See more about Charles H. Sternberg here.

                                                                        Kansas Academy of Science.                                                         119

Explorations of the Permian of Texas and the Chalk of

Kansas, 1918.

CHARLES H. STERNBERG.

     The splendid skeleton of Dimetrodon gigas I collected in 1917 and sold to the United States National Museum has been mounted at last, and is one of the world's famous specimens. I do not know of a more perfect single individual. It came from the famous Craddock quarry discovered by the late Doctor Williston's assistant, Mr. Miller. Through the kindness of Mr. Craddock, the owner of the quarry, I not only collected there in 1917, but last year. Owing to the fact that the quarry is now covered with about twenty feet of earth and clay of the toughest character the work was very difficult and I was obliged to employ a man with a heavy team of horses, with plow and scraper. I succeeded in securing many more or less perfect skeletons of several species. Unfortunately, none were as perfect and capable of making into a fine open mount as the National Museum specimen. This quarry is in the face of a hill. As I have gone deeper and deeper into the hill the manner in which the animals were stranded here becomes more and more apparent. On the very bottom of the quarry are innumerable bones of very small animals, Seymoriana and other batracians, etc. They are usually scattered and are free from matrix, consequently they are among the rarest of Permian vertebrates that have not been injured by the encrusting silica that covers all the other bones at higher levels, and which is so difficult. to remove. The National Museum specimen came from near this level. Above are about four or, five feet in the heavy, fine-jointed red clay. Though water has filtered and coated all the bones with silica, I found several more or less perfect skeletons. Often the entire column, except the tail, with enormous spines were present; sometimes the arches and limbs, as in the best specimen we found, discovered by my son, George F.  The longest spine of this individual was four feet. Most of the column and tail were present. The skull was disarticulated. The arches seemed present. Many of the spines, however, were twisted and interwoven, all the bones covered with a thin coating of silica. As I understood Doctor Gilmore, it took two preparators a year to prepare and mount the National Museum specimen. You will realize something of the labor it will take to prepare this one. This, with my whole collection from the Craddock quarry, I sold to the American Museum.

     From Seymour, Tex., my boys, George and Levi, drove my car to the Rock creek Horse quarry, near Tulia, Tex.. but the formidable mass of sand that lay above it induced them to turn their Ford truck northward, and I joined them in the Kansas chalk in Logon county, on Butte creek. I was so fortunate as to find a fine tylosaur skeleton the second day in the field. There were twenty-one feet of the skeleton present in fine chalk. The complete skull was crushed laterally, nearly the complete front arches and limbs were present, as was also the pelvic bones and both femora. All the vertebræ to well into the caudal region beyond the lateral spines were continuous, with the ribs in the dorsal region. Between the ribs was a large part of a huge plesiosaur with many half-digested bones, including the large humeri, part of the

     120                                                                 Kansas Academy of Science.

 

coracoscapula, phalanges, vertebræ, and, strangest of all, the stomach stones, showing that this huge tylosaur, that was about twenty-nine feet long, had swallowed this plesiosaur in large enough chunks to include the stomach. How powerful the gastric juice that could dissolve these big bones! This specimen I sent to the United States National Museum. A little Clidastes skeleton found by my son Levi proves not only to be new, but possessed of remarkable characters not yet described. I will only in a general way give you an idea of this little sea lizard. It is 8 ½ feet long, skull 14 inches long. Levi in preparing the skeleton restored the maxilla, jugals, nasals and prosquamosals, with the ends of some of the teeth; also four of the cervical and one dorsal vertebra. These had been destroyed by incrusting gypsum. The bifrucated coracoids, 2 inches wide, are large compared to the scapulæ, which are only 1 ¼ inches wide. The most remarkable thing about this mosasaur is the fact that the humeri and femora have distinct, round heads, similar to those of mammals. Further, all the Clidastes humeri I know are broad, square bones, nearly as broad as they are long. In this specimen the humerus is 2 ¼ inches, while it is only 1 ¼ inches wide in the widest part; the same with the femur. The front paddle is 7 inches long to ends of first row of phalanges; width only 2 ½ inches. The column is continuous to the pygals, where they are scattered. The pelvic arches and paddles are only partially preserved. The caudals are beautifully preserved, with a high fin in the last half of the column; the chevron well preserved, with all of them anchylosed to the centra of the vertebræ. The only genus among the mosasaurs where this is the case. They usually are distinct, the proximal heads fitted snugly into little basins hewn out., as it were, from the centra of the vertebræ. I own this new Clidastes. [now Eonatator (Halisaurus) sternbergi, described by Carl Wiman, 1920; revised to Halisaurus by Russell (1967) and most recently to Eonatator (novem gen.) by Bardet, et al., 2005).

    We secured two specimens of Platecarpus coryphæus of exactly the same size. One found by Levi Sternberg has much of the head, the column, and ribs to the pygals. The second, found by George F. Sternberg, has a very complete tail to the very end, and the pelvic arch and one paddle; other specimens furnished the rest of the paddles. We have mounted this as a slab specimen, and makes a skeleton 17 feet long-very impressive, indeed. George F. Sternberg found also a very fine skull and most of the skeleton of a small Pteranodon. The skull is only 27 inches long; missing, only the crest and front of the mandibles. Both coracoscapula are present, with one humerus, radius, ulna, carpal joint., and most of one elongated finger. Both hind limbs and several vertebræ and ribs. This, when prepared, will be one of the few fine skulls of these flying reptiles. We secured also a very fine Portheus, complete, nearly, except the ends of the ribs, the dorsal and caudal fins, and 18 caudal vertebræ. These we have restored, and we have a fish 13 feet long, with spread of tail fins of 35 inches. These are the chief discoveries of my last season's labor in the Permian of Texas and the chalk of Kansas.

 

 

smith-ta.jpg (25397 bytes) LEFT:  The skull of the Tylosaurus proriger (USNM 8898) in the exhibits of the Smithsonian Institution. This skull is approximately 1.2 m (4 ft.) long and may represent the maximum size of Tylosaurus during the deposition of the Smoky Hill Chalk.  The two bones shown under the jaw are part of the hyoid apparatus - rarely preserved in mosasaurs.

LEFT: One of the two propodials (a humerus or femur) recovered as gut contents from inside the rib cage of the Tylosaurus proriger. Note partially digested appearance.

RIGHT: The second of two propodials recovered by Sternberg, even more severely damaged by stomach acids.

LEFT: An assortment of partially digested bone fragments from the plesiosaur.

RIGHT: A ventral view of four caudal vertebrae and three caudal ribs, damaged by stomach acids. Bruce Schumacher (pers. comm. 2004) thinks that these may instead be cervical vertebrae based on the large nutritive foramina. 

LEFT: All twenty-one epipodials (finger bones) recovered by Sternberg from the remains of the plesiosaur.  A single Dolichorhynchops or Polycotylus paddle would have up to 70 epipodials.

RIGHT: An close-up, end view of one of the four caudal vertebrae shown above at right; note damage to the surface of the bone by stomach acids.  

For pictures of the gastric contents of another Tylosaurus proriger (this one from South Dakota), go HERE (middle of page).

JUST PUBLISHED (2004) - READ AN UPDATED VERSION OF THE PAPER HERE

Everhart, M. J. 2004. Plesiosaurs as the food of mosasaurs; new data on the stomach contents of a Tylosaurus proriger (Squamata; Mosasauridae) from the  Niobrara Formation of western Kansas. The Mosasaur 7:41-46.

 

ABSTRACT

Although plesiosaurs and mosasaurs co-existed for about 25 million years at the end of the Cretaceous, the fossil record was mute regarding interactions between these two groups of marine reptiles until a discovery was made in the summer of 1918. At that time, Charles F. Sternberg uncovered the partially digested bones of a plesiosaur as stomach contents in an adult (9 m) Tylosaurus proriger skeleton in the Smoky Hill Chalk Member (early Campanian) of the Niobrara Formation near Twin Butte Creek in Logan County, Kansas. Sternberg reported his discovery at the annual meeting of the Kansas Academy of Science in 1919 and indicated that the material had been sent to the United States National Museum. Due to unusual circumstances regarding the publication of his brief paper in the Transactions of the Kansas Academy of Science, however, the association of the two specimens went largely unnoticed until 2001. This association demonstrates conclusively that mosasaurs fed on plesiosaurs and provides additional data about the ecology of the Western Interior Sea. Here the remains are re-examined and discussed in light of related information that has become available in the more than eighty years since their original discovery.

 

Suggested references:

 

Bardet, N. & X. P. Suberbiola. 2001. The basal mosasaurid Halisaurus sternbergii from the Late Cretaceous of Kansas (North America): a review of the Uppsala type specimen. Comptes Rendus de l'Académie des Sciences - Series IIA - Earth and Planetary Science 332:395-402.

 

Bardet N., X. P. Suberbiola, M. Iarochene, F. Bouyahyaoui, B. Bouya, and M. Amaghzaz. 2005. A new species of Halisaurus from the Late Cretaceous phosphates of Morocco, and the phylogenetical relationships of the Halisaurinae (Squamata: Mosasauridae). Zoological Journal of the Linnean Society 143: 447-472.

 

Everhart, M. J. 2004. Plesiosaurs as the food of mosasaurs; new data on the stomach contents of a Tylosaurus proriger (Squamata; Mosasauridae) from the  Niobrara Formation of western Kansas. The Mosasaur 7:41-46.

 

Gilmore, C. W. 1921. An extinct sea-lizard from western Kansas. Scientific American, CXXIV 273, 280, 3 figs.

 

Martin, J. E. and P. R. Bjork, 1987. Gastric residues associated with a mosasaur from the late Cretaceous (Campanian) Pierre Shale in South Dakota, Dakoterra, 3:68-72.

 

Sternberg, C. H. 1922. Explorations of the Permian of Texas and the chalk of Kansas, 1918. Kansas Academy of Science, Transactions 30(1):119-120.

 

Varricchio, D. J. 2001. Gut contents from a Cretaceous Tyrannosaurid; Implications for Theropod dinosaur digestive tracts. Journal of Paleontology 75(2):401-406.

 

Wiman, C. J. 1920. Some reptiles from the Niobrara group in Kansas. Bulletin of the Geological Institute, Uppsala 18:9-18, 9 figs., pls. II-IV.